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Food-Transport-Phloem | uksir-notes |Plant- Physiology

 Plant Physiology:

 

transport


Want to know about transport of Transpiration, stomata opening and closing, guttation in plants, Click the Link below:


Want to know about transport of water, osmosis in plants, Click the Link below:


Transport of food:

-          Transport of food from source/ production center to the sink/ consumption center is called as transport of food in plants.

-          Leaves are the food production center/ unit in plants, some time young stem may contain chlorophyll and perform photosynthesis.

-          Consumption centers are mostly other parts like stem, apices, root, fruits etc.

-           The transporting tissue is phloem.

-          Phloem sap transport mostly water and carbohydrate.

-          90% of the solute is sucrose, as it is inactive highly soluble.

-          Later sucrose converted to monosaccharide.

-          Phloem may transport- some minerals, amino acids, sugar phosphate, potassium and phosphate ions etc.

-          It can also carry vitamins and phytohormones.

Phloem


Routs of Translocation:

-          Solutes can be transported in various directions.

(a) Downward translocation:

-          Its the most common pathway.

-          Here food translocated from leaves to root and stem in downward movement.

-           Food is utilized in growth and storage.

(b) Upward Translocation:

-          In special condition food condition organic solutes and mineral salts transported in upward direction.

-          Here from root storage transportation occurs to leaves as well as apex.

-          During early stage of development, leaves are non functional. At that time they need food.

-          Thus upward transport occurs.

(c) Lateral Translocation:

-           In woody stem lateral translocation of food occurs.

-          It’s a very slow process.

-          Here food is supplied to the cortex and epidermal layer.

Translocation of plant


Mechanism of Translocation:

-          Several theories have been put forwarded to explain the solute movement:

-          (a) Cytoplasmic streaming:

-          Proposed by Curtis in 1923.

-          Cytoplasm of living cell show streaming movement called as cyclosis.

-          The cytoplasm transport organic food towards all direction.

-          But later it was rejected by Mason and Phyllis.

-          (b) Diffusion Theory:

-          Proposed by Mason and Maskell in 1928.

-          Due to diffusion of solute from higher conc. to lower conc.

-          Later this theory was modified to activated diffusion theory.

Translocation


-           

-          (c) Trans cellular streaming Theory:

-          Proposed by Thaine in 1962.

-          According to him, sieve tubes contain proteinaceous tubular strands.

-          These strands transport solute .

-          (d) Mass flow theory/ Pressure flow theory: (Munch Hypothesis)

-           It was proposed by E. Munch in 1930.

-          According to this theory organic solutes are translocated by phloem sieve tubes.

-           The leaves are the sources where as other parts are the sink.

-           The movement of solutes can be upward or downward, that is bidirectional.

Pressure flow hypothesis


Phloem sap move by following steps:

1. Source: the part of plant where food is prepared.

2. Phloem Loading: during photosynthesis, mesophyll cells produce sugar.

-           This sugar dissolves in cell sap and increase the conc.

-          Thus due to high osmotic pressure and turgor pressure, dissolve sugar transported to companion cell of phloem tissue.

-          Later solutes are transferred to sieve tubes by active transport.

-          This is called as phloem loading.

3. Transport of solutes: in side sieve tube, pressure difference occurs.

-          Pressure is highest at source and lowest at sink.

-          Thus solutes are transported throughout the plant.

-          Necessary water is absorbed from xylem.

4. Sinks : they can be any part, where food can be used.

-          Example: Stem, root, fruit and any storage organ.

5. Phloem unloading: at sink osmotic pressure and TP is always low.

-          Here also solutes are unloaded by active transport.

source to sink



Want to know about transport of Transpiration, stomata opening and closing, guttation in plants, Click the Link below:


Want to know about transport of water, osmosis in plants, Click the Link below:



    

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